Astragalus duchesnensis
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Title
Astragalus duchesnensis
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Author(s)
Rupert C. Barneby
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Scientific Name
Astragalus duchesnensis M.E.Jones
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Description
59. Astragalus duchesnensis
Wiry, sparsely leafy, strigulose throughout or nearly so with fine, straight, appressed (sometimes a few narrowly ascending) hairs up to 0.3-0.55 mm. long, the stems and herbage greenish or cinereous, the leaflets either pubescent on both sides or commonly glabrescent above; stems few or several, sometimes solitary, erect and ascending in clumps or sometimes diffuse and straggling, 1.5—3.5 dm. long, subterranean for a space of 1.5—6 cm., leafless and purple-tinged toward the base, branched or spurred at 1—5 nodes preceding the first peduncle, the branches sometimes again branched, becoming zigzag distally; stipules dimorphic, the lowest ones ovate-oblong, papery-membranous, pallid or brownish, several-nerved, 3-8 mm. long, adnate to the vestigial petiole and amplexicaul-decurrent around 3/4 to the whole stem’s circumference, free, the median and upper ones smaller, triangular or triangular-acuminate, the firm, green or purplish blades erect or recurved; leaves (1.5) 2.5-9.5 cm. long, shortly petioled, with 7-11 (15) rather distant, linear, linear-filiform, or (in some lower leaves) narrowly oblanceolate, obtuse or retuse, involute or rarely flat, petiolulate leaflets 3—20 mm. long, but the leaflets of the uppermost leaves often reduced in size, or number, or absent, the terminal one represented by a scarcely expanded, often falcate blade continuous with the rachis; peduncles erect or incurved-ascending, (3) 4-9.5 cm. long; racemes loosely (3) 6-18 (22)-flowered, the axis becoming (1) 2.5-12 cm. long in fruit; bracts papery or early becoming so, triangular-ovate or lance-acuminate, 0.7-2 mm. long; pedicels at anthesis straight, ascending, 0.8-1.4 mm. long, in fruit arched outward or abruptly downward, thickened, 1-2.2 mm. long; bracteoles 0; calyx 3.6-5 mm. long, strigulose with black or mixed black and white hairs, the oblique or symmetric disc 0.7-1.2 mm. deep, the campanulate, often purplish tube 3.1-4.3 mm. long, 2.3-2.9 mm. in diameter, the broadly triangular-subulate or deltoid teeth 0.4-1 mm. long; petals bicolored, the pale-eyed banner and keel reddish-pink or -purple, the wings white; banner ovate-cuneate, shallowly emarginate, 8.5-12.4 mm. long, 5.2-7.8 mm. wide; wings (about as long or a trifle longer) 8.4-12.5 mm. long, the claws 4-5.2 mm., the oblong-oblanceolate, obtuse, slightly incurved blades 5-8.6 mm. long, 1.8-2.8 mm. wide; keel 8-9.8 mm. long, the claws 3.6-4.8 mm., the half-obovate blades 4.5-5.9 mm. long, 2.4-3.1 mm. wide, incurved through 90-95° to the broadly deltoid, obtuse apex; anthers 0.4-0.75 mm. long; pod deflexed, sessile or attenuate at base into a thick, glabrous, stipelike neck up to 1 mm. long, the body linear or commonly linear-oblanceolate in profile, 2-3.5 cm. long, 3.3-5.3 mm. in diameter, straight or more often gently arched downward in the lower 1/3 and straight or nearly so thereafter, tapering into the narrow base, triangular-cuspidate at apex, dorsiventrally compressed in the lower 1/2 and there obscurely trigonous, with flattened or openly and shallowly grooved dorsal and low-convex lateral faces, becoming ± laterally compressed and 2-sided distally, carinate ventrally by the prominent suture, the thinly fleshy, pale green or purplish, densely strigulose valves becoming stiffly papery and stramineous; ovules 21-31; seeds olivaceous or drab, closely pitted, 2.1-3 mm. long.—Collections: 12 (v); representative: Ripley & Barneby 8714 (CAS, RSA, UTC); W. A. Weber 5313 (COLO, RSA); Rollins 1743 (NY); Barneby 12,706 (CAS, NY, RSA).
Desert flats, hillsides, badlands, dunes, and sandy pockets about rock outcrops, on sandstone, 4800-5100 feet, locally plentiful on the floor of the central Uinta Basin, from the lower Duchesne River east to Dinosaur National Monument and to the White River a few miles west of the Colorado State line, in Uintah and adjoining Duchesne Counties, Utah.—Map No. 24 (some stations after Graham, 1937, p. 247).—Late April to June.
Astragalus duchesnensis (of Duchesne River or County) Jones, Contrib. West. Bot. 13: 6. 1910 ("Duchesnensis").—"...from 13 miles below Theodore Utah to Chepeta Well and White River near the Colorado line... ’’—Lectotypus, collected at Chepeta Well, "6000 ft.," May 23, 1908, Jones, POM (3 sheets, marked "type" in Jones’s hand).—Paratypi, collected by Jones between Theodore and Myton, 5000 ft., May 19, 1908, CAS, DS, NY (2 sheets), POM (5 sheets), RM, US!—Lonchophaca duchesnensis (Jones) Rydb. in N. Amer. Fl. 24: 313. 1929.
The Duchesne milk-vetch is a thinly leafy plant, but hardly junceous, since the foliage is better developed than in most other Lancearii, with only some uppermost leaves reduced (or nearly so) to the naked rachis. The ascending, bicolored flowers, unusual for the reddish, distinctly vinous tint of the banner and keel, and the curiously compressed fruits are distinctive. The known area of dispersal extends not more than thirty-five miles east and west over the floor of the Uintah Basin, but the species is not uncommon in its region. The plants occur in colonies and in favorable years form bold clumps of stems, especially vigorous on sands of the red beds but equally common on the whitish or tawny sand-clays of the badlands; however in dry years a formerly flourishing colony may yield only a few straggling wisps of stem, sterile or nearly so.
According to Jones s last account (1923, p. 251, Pl. 64) the pod of A. duchesnensis may reach a length of 6 cm. and a width of 7 mm., substantially larger than anything found in the Jones herbarium or in any other collection studied.