Marina pueblensis
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Title
Marina pueblensis
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Author(s)
Rupert C. Barneby
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Scientific Name
Marina pueblensis (Brandegee) Barneby
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Description
22. Marina pueblensis (Brandegee) Barneby
(Plate XVIII)
Slender annuals from an orange taproot, branching from near the base or from well below the middle, 1-5.5 (6) dm tall, appearing glabrous to the inflorescence (the leaflets really so), but the leaf-stalks, the stems (minutely punctate but not verruculose) at least distally, and the raceme-axis finely pilosulous with subappressed or spreading-incurved hairs up to 0.15-0.25 mm long, the foliage green, the leaflets lineo- late both sides, smooth above, paler and minutely punctate beneath, their often purple margins gland-crenulate; leaf-spurs less than 0.5 mm long; stipules narrowly subulate to linear-attenuate, 0.8-2 mm long, purple-castaneous becoming dry and fragile; intrapetiolular gland small; post-pedolular glands prominent, subulate, orange; main cauline leaves 1.5-3.5 cm long, shortly petioled, with scarcely margined rachis and (2) 3-5 pairs of oblanceolate, oblong-elliptic, or obovate, subacute or obtuse and gland-mucronate, flat leaflets 2.5-6.5 mm long, the terminal leaflet similar but longer, mostly 5-11 mm long, stalked beyond the last pair; peduncles very slender, mostly leaf-opposed, (0.4) 1-3 cm long; racemes loosely or remotely (4 ) 8-20 ( 25)-flowered, the flowers ascending, the axis (1) 2-9 cm long; bracts ovate-acuminate, sheathing at base, 0.8-2 mm long, persistent, early becoming brown and glumaceous, glabrous dorsally, minutely gland-fimbriolate; pedicels subfiliform, 0.9-1.4 mm long, charged at apex with a pair of minute glands; calyx 3-3.7 mm long, thinly pectinate-pilosulous especially along the ribs with stiff, spreading-incurved, lustrous hairs up to 0.15-0.25 mm long, the tube accordion-pleated, (1.5) 1.7-2 mm long, its orifice subsymmetrical, the recessed, membranous intervals charged with a chain of 5-8 small, yellowish glands, the membranes produced high into the teeth, the teeth unequal, the dorsal one longest and proportionately narrowest, 1.3-1.9 mm long, the ventral and lateral pairs shorter, obovate-short-acuminate, 1.1-1.5 mm long; petals vivid magenta-purple except for a pale (rubescent) eye-spot on blade of banner, all eglandular, the inner ones perched at 1.2-1.5 mm above hypanthium rim, the keel-blades united by narrow overlap; banner 1.7-2.4 mm long, the claw 1.1-1.6 mm, the reniform or angled, emarginate or subacute, incipiently peltate blade 1-1.4 mm long, 1.3-1.8 mm wide; wings 2.3-2.9 mm long, the claw 0.6-0.8 mm, the obovate-oblong blade 1.8-2.3 mm long, 1-1.5 mm wide; keel 2.5-3.4 mm long, the claws 0.7-1.3 mm, the obovate blades 1.8-2.4 mm long, 1.2-2.6 mm wide; androecium 9 (rarely 10)-merous, (2.8) 3.3-4 mm long, the longer filaments free for 0.8-1 mm, the connective gland-tipped, the pale blue anthers 0.3-0.7 mm long; pod 2-2.4 mm long, obovate-harp-shaped in profile, the style-base terminal but excentric, sharply prowed, the valves membranous at extreme base, thence papery, thinly pilosulous, and charged with 2 crescents of blister-glands; seed ±1.6 mm long; 2n = 20 (Mosquin).— Collections: 8 (v).
Dry stony or grassy hillsides, often in drought-deciduous thorn-scrub, on calcareous, volcanic, and perhaps other bedrock, 855-1200 m (± 2850-5750 ft), local but forming colonies, e. arm of Balsas Depression in Morelos, Puebla, and n. Oaxaca, extending e. in Puebla across the low divide to the headwaters of Rio Salado in the cuenca de Rio Papaloapan. — Flowering September to November. —Material. Morelos. Cuernavaca, Matuda 222 (MEXU); Tlatizapan: near Moyotepec, Ripley & Barneby 13,702 (CAS, NY, US). Puebla. Tehuacan: Purpus 5649 (F, UC); Coxcatlan (typus); Izucar de Matamoros: 8 mi. s.-e. of Izucar, Ripley & Barneby 13,689 (CAS, GH, K, MEXU, MICH, NY, US); Tehuitzingo: Tehuitzingo, Ripley & Barneby 14,575 (DAO, IA, NY, US); Petlalcingo: n.-w. of Petlalcingo, Ripley & Barneby 14,576 (CAS, DAO, NY, US). Oaxaca. Huajuapan de Leon: Chazumba, near the Puebla boundary, Ripley & Barneby 14,706 (NY).
Marina pueblensis (Brandg.) Barneby, comb. nov., based on Dalea pueblensis (of Estado de Puebla) Brandg., Univ. Calif. Pub. Bot. 4: 86. 1910. — "Collected near Coxcatlan, No. 4146..." — Holotypus, collected in Sept 1909, by Carl Anton Purpus, UC! isotypi, F, GH, NY, US! — Parosela pueblensis (Brandg.) Rydb., N. Amer. FI. 24: 52. 1919.
This is a thin-stemmed, sparsely leafy plant, not easily singled out from the tangle of grasses and other small herbs among which it delights to grow, except early in the day, before the caducous, small but vivid magenta-purple petals have fallen to the ground. The species is strongly characterized by the annual root, shining, wiry stems, few (mostly 7-11) leaflets ample in context of ser. Unifoliolatae, and openly or remotely flowered racemes of tiny, pectinately ciliolate calyces, each subtended by a little persistent, papery brown bract. According to the label the type collection came from elevations of 7000 - 8000 ft, but this is probably a mistake on Brandegee’s part. Coxcatlan lies on the floor of Rio Salado valley downstream from Tehuacan, and all subsequent collections of M. pueblensis have been made at levels far below the 2000 meter contour.
For discussion of kinship, see under M. melilotina (above) and 24. M. unifoliolata.