Thelypteris biolleyi (Christ) Proctor

  • Authority

    Mickel, John T. & Smith, Alan R. 2004. The pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1-1054.

  • Family

    Thelypteridaceae

  • Scientific Name

    Thelypteris biolleyi (Christ) Proctor

  • Description

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    Species Description - Rhizomes short-creeping to suberect; fronds monomorphic, or nearly so; stipes to 50 cm long or more, brownish, stellate-hairy; blades dark green, herbaceous, pustulate, to 75 cm long with pinnatifid apices; pinnae 10-20 pairs, 12-22(-28) x 2-3(-4) cm, sessile or short-stalked to 1 mm, incised ca. 0.5-0.7 the way to the costae, proximal pinnae often narrowed at their bases; segments suboblique, subfalcate, rounded at apices, ca. 4-6 mm wide; veins to 16 pairs per segment, the lowermost pair from adjacent segments anastomosing at a 60-90º angle or connivent at sinuses; indument abaxially of numerous anchor-shaped (2 recurved branches at tips) hairs 0.2-0.5 mm long on costae, costules, and tissue between veins, also with a few sessile, appressed stellate hairs (4 arms), adaxially the tissue between veins with numerous appressed stellate hairs; sori inframedial to medial, exindusiate or indusiate; sporangia glabrous or with a few minute stellate hairs; 2n =72 (Jam).

  • Discussion

    Aspidium biolleyi Christ in Pittier, Prim. Fl. Costaric. 3(1): 31. 1901. Dryopteris nephrodioides (Klotzsch) Hieron. var. biolleyi (Christ) C. Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 248. 1913. Goniopteris biolleyi (Christ) Pic. Serm., Webbia 31: 251. 1977. Type. Costa Rica. Tuis pre`s Turrialba, Pittier 11243 (P?; isotypes CR, US!). Complete synonymy given in Smith (1983).

    Thelypteris biolleyi differs from all other Mexican species of Goniopteris except T. hatchii in having sessile, appressed, stellate hairs on both sides of the blades, between the veins. From T. hatchii it differs in having some anchor-shaped hairs on the costae and laminae abaxially. The two Oaxacan collections have relatively large indusia and proximal pairs of veins connivent at sinuses. In these two characters they match better T. nephrodioides (Klotzsch) Proctor from the Antilles and South America. However, they bear abundant anchor-shaped hairs characteristic of T. biolleyi.

  • Distribution

    In tropical and montane rain forests; 250-1150 (-1600) m. Mexico; Guat, Bel, Hond, Nic, CR, Pan; Jam; Col, Ven, Fr Gui, Ec, Peru, n Braz.

    México Mexico North America| Guatemala Central America| Belize Central America| Honduras Central America| Nicaragua Central America| Puntarenas Costa Rica Central America| Jamaica South America| Amazonas Colombia South America| Venezuela South America| Approuague-Kaw French Guiana South America| Amazonas Ecuador South America| Peru South America| Brazil South America|