Arachniodes denticulata (Sw.) Ching

  • Authority

    Mickel, John T. & Smith, Alan R. 2004. The pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1-1054.

  • Family

    Dryopteridaceae

  • Scientific Name

    Arachniodes denticulata (Sw.) Ching

  • Description

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    Species Description - Rhizomes ascending to suberect, short, compact; rhizome scales brown, lustrous, 10–20 x 1–2 mm, linear-lanceolate, entire; fronds clustered, mostly (30–)50–90 cm long; stipes castaneous at base, often stramineous distally, 12–32 cm x (1–)2–4 mm, 1/3–1/2(–2/3) the frond length, glabrous except for linear brown scales at bases; blades dark green adaxially, lighter green abaxially, chartaceous to subcoriaceous, 12–30 x 8–26 cm, deltate to pentagonal, 4-pinnate at bases, the proximal pair of pinnae strongly inequilateral, greatly exaggerated basiscopically, anadromous; rachises glabrous; pinnae alternate; segments oblanceolate to obovate, 5–8 x 3–4 mm, bases cuneate, margins strongly dentate with spinulose tips; indument of costae and costules of sparse, hair-like, fibrillose scales; indusia tan to red-brown, glabrous, persistent, 1–2(–3) mm diam.; 2n=82 (Oax, Chis), 164 (Jam).

  • Discussion

    Polypodium denticulatum Sw., Prodr. 134. 1788. Aspidium denticulatum (Sw.) Sw., J. Bot. (Schrader) 1800(2): 40. 1801. Polystichum denticulatum (Sw.) J. Sm., J. Bot. 4: 195. 1841. Dryopteris denticulata (Sw.) Kuntze, Revis. Gen. Pl. 2: 812. 1891. Rumohra denticulata (Sw.) Copel., Gen. Fil. 114. 1947. Byrsopteris denticulata (Sw.) C. V. Morton, Amer. Fern J. 50: 152. 1960. Type. Jamaica. Swartz s.n. (S; isotype C).

    Aspidium jucundum Fe´e, Me´m. Foug. 10: 41. 1865. Dryopteris denticulata f. jucunda (Fe´e) C. Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 115. 1920. Lectotype (chosen by Christensen, 1920: 115). Mexico. [Oaxaca:] Galeotti 6563 (P; isotypes BR-photos UC!, US!, RB).

    Christensen (1920) recognized eight named forms and varieties and several unnamed ones of this polymorphic species. These variants differ chiefly in size and dissection of the blades, blade texture, and other rather nebulous characteristics. Chromosome counts indicate that polyploidy is a part of this variability. Christensen’s forms seem to be connected by numerous intermediates, so we choose not to apply infraspecific names to Mexican material.

  • Distribution

    Terrestrial in wet montane forests; 1000–2900 m. Mexico; Guat, Bel, Hond, Salv, Nic, CR, Pan; Cuba, Jam; Col, Ven, Guy, Ec, Peru, Braz, Bol.

    Bolivia South America| Brazil South America| Peru South America| Ecuador South America| Guyana South America| Venezuela South America| Jamaica South America| Cuba South America| Panama Central America| Costa Rica South America| Nicaragua Central America| El Salvador Central America| Honduras Central America| Belize Central America| Guatemala Central America| México Mexico North America|