Pseudo-calliergon trifarium (F.Weber & D.Mohr) Loeske

Authority

Hedenäs, Lars. 2003. Amblystegiaceae. Fl. Neotrop. Monogr. 89: 1--107. (Published by NYBG Press)

Family

Campyliaceae

Scientific Name

Pseudo-calliergon trifarium (F.Weber & D.Mohr) Loeske

Type

Type. Sweden. Uppland: Uppsala, (probably) F Weber & D. Mohr s.n. (lectotype, B, designated by Hedenas, 1989a; probable isolectotype, C).

Synonyms

Hypnum trifarium F.Weber & D.Mohr, Calliergon trifarium (F.Weber & D.Mohr) Kindb., Scorpidium trifarium (F.Weber & D.Mohr) H.K.G.Paul

Description

Species Description - Plants medium-sized, rarely larger, slightly turgid; green, brownish green, yellowish, or yellow brown, rarely with a golden gloss when dry. Stem unbranched or slightly and irregularly branched, somewhat stiff; pseudoparaphyllia broad; axillary hairs with 1-2-celled upper part, this early yellowish (rarely hyaline), often with elongate apical cell. Stem leaves straight, erect and imbricate or sometimes spreading, broadly or very broadly ovate, not plicate, concave or strongly concave; apex ± broadly rounded or occasionally broadly obtuse; margin entire; costa single, extending 60-90(-95)% of way up leaf; median laminal cells (30.5-)33.0-104.0(-107.0) × 5.0-11.0 µm, incrassate and porose, rarely thin-walled and eporose; alar cells long-rectangular, inflated, incrassate or strongly incrassate (except when young), somewhat porose and with yellow walls when mature; alar group approximately transverse-triangular or less commonly cells in single transverse row, indistinctly delimited from surrounding cells; supra-alar cells quadrate to rectangular, in four or more rows of cells along leaf margin above alar cells, indistinctly delimited from surrounding cells and sometimes impossible to delimit from both alar cells and laminal cells farther up. Inner perichaetial leaves narrowing ± abruptly to obtuse or acute apex. Lower outside of exostome cross-striolate. [Sporophytes not known from neotropical material.]

Discussion

Pseudo-calliergon trifarium was known from Venezuela and the Dominican Republic (Buck, 1998; Delgadillo et al., 1995), and is here reported from Colombia as well. This species is usually easily recognized by its stiff, unbranched or almost unbranched shoots with leaves at least partially closely imbricate. The stem leaves are broadly or very broadly ovate; the leaf apex is usually broadly rounded, occasionally broadly obtuse; and the alar cells form a transverse-triangular group (occasionally a single row of cells) and extend far inward from the leaf margin. Pseudocalliergon trifarium is sometimes confused with Straminergon stramineum, but S. stramineum is whitish or pale (vs. a dark yellow to yellow brown secondary coloration); has mostly narrower (oblong to narrow-ovate) leaves; and has ovate alar groups along the basal margin of the leaf.

Distribution and Ecology: Dominican Republic (2200 m a.s.l.), Venezuela (3800-3980 m), and Colombia (3800-3850 m). Widespread in N America and Eurasia, where it is most common in boreal areas, but occurring in both the arctic and temperate zones. In S America known from brook and lake shores, and submerged in a lake; outside this area also in deep fens. In other areas of the world this species occurs in relatively mineral-rich habitats, and this is most likely the case at its neotropical localities.

Distribution

Dominican Republic South America| La Vega Dominican Republic South America| Venezuela South America| Mérida Venezuela South America| Colombia South America| Boyacá Colombia South America| Cundinamarca Colombia South America|